CONCEPCIÓN, Bolivia — The aging musical score wasn’t easy to read. It was a copy of a copy of a Latin Mass by the 18th-century composer Domenico Zipoli that had crossed the Atlantic and most of South America, only to be stuffed into a box for three centuries in a derelict jungle church where humidity had taken its toll.
And then there were the termites.
The insects had eaten large tracts of the Mass, including the 22nd and 23rd measures.
But while much of the work of Zipoli has vanished in his native Europe, it has managed to survive in eastern Bolivia — along with his vast Baroque musical tradition, which hums through the tropical lowlands.
Here near the borders of Brazil and Paraguay, harpsichords and lutes can be found in the smallest villages. Luthiers have carved violins from local cedar for centuries.
And troves of ancient manuscripts, more recently rediscovered in church archives, have once again revived Zipoli and other composers of the period, whose music is played in elementary schools and on the radio.
“The Baroque is our tradition here,” said Juan Vaca, an archivist in Concepción, leafing through the crumbling pages of the Zipoli Mass with a pair of gloves and a small rod.
The score is a legacy of Jesuit missionaries who left a musical time capsule in Bolivia. By the 1700s, parts of what is now Paraguay, eastern Bolivia and southern Brazil were vast forests of seminomadic native peoples and the slave traders who hunted them. Surrounding the jungles were the Spanish and Portuguese Empires.
The Jesuits descended into the jungles with the twin goals of converting indigenous tribes and sheltering them from enslavement. In the process, they formed a state-within-a-state ruled by priests and local chieftains.
This obscure corner of Latin American history had a brief moment in the Hollywood spotlight with the release of the 1986 film “The Mission,” starring Robert De Niro.
“It was about building a different society, a kind of utopia with education, self-sustainability — and of course, with music, which was the way the Jesuits evangelized,” said the Rev. Piotr Nawrot, a Roman Catholic priest from Poland who lives in Bolivia and was involved in recovering some of the original Baroque manuscripts.
Overall, the Catholic Church’s record in the area was mixed; it agreed to force indigenous groups off many missions they had constructed to solve a territorial dispute between Spain and Portugal. Refusing to leave, some of the indigenous people fought a bloody war, and many of the churches fell into ruin.
But among the lowland Bolivians, the legacy of Baroque music survived — even centuries after indigenous communities lost the tradition of reading music, learning songs by ear.
To understand just how powerful the tradition remains today, consider Urubichá, a farming village northwest of Concepción on the end of an unpaved dirt road bordering a swamp reached only after crossing ten bridges through dense jungle.
The town of 8,000 has a music school teaching 500 students, nearly every child there. At lunchtime, children wander the village square toting instrument cases on their backs. They speak Guarayo, the native language.
“The Guarayo live with this music in their souls,” said Leidy Campos, 32, who teaches music in the village. “People say they were born with a violin in their hands.”
Across a field from the classrooms, Ideberto Armoye, a carpentry instructor, stood in a workshop of half-built violas and violins made from local cedar and mahogany. They were the only woods that could withstand the tropical heat, he said.
To prove his point, he pulled out a violin that had come recently from a factory in China.
“Anything happens to this instrument, look at this big crack,” he said.
While many of the era’s songs had been passed down orally through families in Bolivia, the orchestrations and choral works were thought to be lost. For years it remained one of the mysteries of the time: While Baroque music had been the bridge between the Jesuits and Bolivians, no one knew exactly how it sounded.
“I had to make a deep mental effort to imagine what it might have been like,” said Ennio Morricone, who composed the soundtrack to “The Mission” in the years before the manuscripts were discovered, using a combination of European and indigenous influences.
In the 1990s, Father Nawrot went looking for what might have remained of the written music, a search that led him to the Moxos area further west. He asked village elders in the area about manuscripts of that time. But first, he said, they had questions for him.
“For three hours they questioned me about my faith, my religion,” Father Nawrot recalled. “It was a complete reversal of roles.”
Eventually the Moxos leaders revealed something that astounded him. Thousands of pages of manuscripts, including music from baroque operas to concertos for solo instruments, some of which had been copied as late as 2005, had survived.
The copyists had even signed some of the scores “maestro capilla,” a Baroque-era title used by composers like Johann Sebastian Bach.
“The manuscript was never lost, we just didn’t know about it,” the priest said.
Through much of the 1990s, Father Nawrot worked with Mr. Vaca, the archivist in Concepción, to reassemble another collection of scores that had been found in the 1970s, including the Zipoli manuscripts that had been eaten by termites.
That body of work, which includes both copies of known works as well as unknown pieces written in Bolivia, is now known in classical music circles as Mission Baroque.
The music has admirers well beyond the Bolivian lowlands. One is Ashley Solomon, a professor at the Royal College of Music in London who traveled to the city of Santa Cruz this April to conduct at a festival of baroque music held once each two years at the former Jesuit missions.
“They took this music and made it their own — it’s more upbeat, more positive,” Mr. Solomon said. “The music lifts the soul, rather than self-flagellates, which is what you see in a lot of Western Classical music of the same time.”
And the pieces are shorter, Mr. Solomon said, written in bite-sized increments that more easily hold attention, which tends to wander more now than it did then.
On a recent night not long after sunset, César Cara, the academic director at the music school in Urubichá, led his student orchestra through a rehearsal of “Sonata XVIII,” a score by an anonymous composer who wrote it somewhere in the surrounding hills in the 18th century.
A large insect crawled across the floor as the chorus waited their turn. One of the sopranos squashed it with her foot and kicked it toward the violins.
“We want people to applaud for us because of our level,” Mr. Cara said, noting that the students had recently played with a visiting group from the Juilliard School for one of the concerts in the festival.
Mr. Solomon, the British musician, said the talent was strong in Bolivia. But he also said the villagers had a connection to the music that is rare in Europe, where classical music tends to live apart from popular culture.
Mr. Solomon recalled years ago giving a concert in San Javier, west of Concepción and the site of a sprawling white-and-black Jesuit mission whose wood facade overlooks the main square.
When his group, Florilegium, began to play an 18th-century flute concerto, “Pastoreta Ychepe Flauta,” he was amazed, he said, to hear members of the audience, townspeople who knew the piece, humming the music too.
“We could play Vivaldi’s ‘Four Seasons’ in London and no one would be singing along,” Mr. Solomon said. “But in Bolivia people took the music for their own — and got to the core essence of what music is about.”